Research led by Chinese scientists has unveiled a unique deepwater fish, Branchiostegus sanae, drawing inspiration from the beloved character San of "Princess Mononoke," illustrating the intersection of pop culture and marine biology.
A Deepwater Fish Named After Princess Mononoke's San

A Deepwater Fish Named After Princess Mononoke's San
A new fish species has been named in honor of the iconic warrior princess from Studio Ghibli, inspired by its striking features.
The newly identified species, Branchiostegus sanae, represents a significant contribution to marine biodiversity and pays homage to a powerful environmental narrative.
A striking visual connection between a newly discovered deepwater fish and the warrior princess San from the iconic Studio Ghibli film "Princess Mononoke" has been made, resulting in the fish being named Branchiostegus sanae. Researchers from the South China Sea Institute of Oceanology and other academic institutions identified the fish, noting its unique cheek stripes that resemble San’s facial markings.
In "Princess Mononoke," San is a fierce warrior raised by wolves and fights passionately to protect her forest home, embodying themes of harmony between humanity and nature. The film, a phenomenal success since its release in 1997, remains a cultural touchstone, grossing over 19 billion yen in Japan alone.
The new species belongs to the family Branchiostegidae, well-known for their deepwater habitats that reach depths of 600 meters. Although the diversity of deepwater tilefish is relatively low, with only 31 species described in their family and 19 in their genus, the discovery of Branchiostegus sanae is particularly notable. Lead researcher Haochen Huang highlighted the importance of their findings, remarking on the rarity of discovering a new species in this group—only three new members of the genus have been found in the past 34 years.
The specimens of this newly discovered fish are now preserved in marine biological collections, allowing for further study and insights into their ecological roles. The study not only adds to our understanding of underwater biodiversity but also serves as a reminder of the interconnectedness of culture, conservation, and our natural world.
A striking visual connection between a newly discovered deepwater fish and the warrior princess San from the iconic Studio Ghibli film "Princess Mononoke" has been made, resulting in the fish being named Branchiostegus sanae. Researchers from the South China Sea Institute of Oceanology and other academic institutions identified the fish, noting its unique cheek stripes that resemble San’s facial markings.
In "Princess Mononoke," San is a fierce warrior raised by wolves and fights passionately to protect her forest home, embodying themes of harmony between humanity and nature. The film, a phenomenal success since its release in 1997, remains a cultural touchstone, grossing over 19 billion yen in Japan alone.
The new species belongs to the family Branchiostegidae, well-known for their deepwater habitats that reach depths of 600 meters. Although the diversity of deepwater tilefish is relatively low, with only 31 species described in their family and 19 in their genus, the discovery of Branchiostegus sanae is particularly notable. Lead researcher Haochen Huang highlighted the importance of their findings, remarking on the rarity of discovering a new species in this group—only three new members of the genus have been found in the past 34 years.
The specimens of this newly discovered fish are now preserved in marine biological collections, allowing for further study and insights into their ecological roles. The study not only adds to our understanding of underwater biodiversity but also serves as a reminder of the interconnectedness of culture, conservation, and our natural world.